Australopithecus africanus was an early homini, an australopithecine, who lived between 2-3 million years ago in the Pliocene. In common with the older Australopithecus afarensis, A. africanus was slenderly built, or gracile, and was thought to have been a direct ancestor of modern humans. Fossil remains indicate that A. africanus was significantly more like modern humans than A. afarensis, with a more human-like cranium permitting a larger brain and more humanoid facial features. A. africanus has been found at only four sites in southern Africa - Taung (1924), Sterkfontein (1935), Makapansgat (1948) and Gladysvale (1992).